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DiversiTree: diversifying our woodlands to increase resilience

Lead Research Organisation: James Hutton Institute
Department Name: Ecological Sciences

Abstract

The UK government plans to increase woodland cover as part of its plans to store more carbon, to mitigate climate change. However, many of the UK's trees are threatened by climate change and a range of pests and diseases, which might limit their ability to contribute to carbon storage and the wide range of other benefits delivered by woodlands. We therefore need to make our woodlands resilient to these future threats. Resilience is the ability of a system, such as a woodland, to recover from a disturbance. One commonly proposed approach to increase the resilience of woods is to increase their tree diversity. Thus, spreading the risk amongst many different trees, as we don't know exactly how each tree species will respond to climate change, nor what threats from pests and diseases they may face decades into the future. However, woodland managers have different perceptions of diversity, and how management may best deliver it, and we know that different tree species will support the woodland ecosystem in different ways. Therefore, it is important to combine stakeholders' knowledge with ecological knowledge to identify which tree species and management approaches best deliver diversification that increases resilience.

DiversiTree focuses on woods dominated by two conifer species, Scots Pine and Sitka Spruce, as in the year to March 2021 54% of all new woodland was coniferous. Scots Pine is the UK's only native conifer of economic significance. It is planted for timber production but is also the dominant species in the culturally iconic native Caledonian pinewoods. Scots Pine is at risk from the tree disease Dothistroma. Sitka Spruce is not native to Britain but is our most economically valuable tree species and is at risk from invasive bark beetles and climate change.

This project addresses four knowledge gaps related to the diversification of woodlands: 1) How do stakeholders understand forest diversity, their diversification strategies, and their visions and ambitions for diverse future forests? 2) Are the microbes found on the leaves of trees more diverse in woodlands with mixed tree species and does this help trees to better defend themselves against diseases? 3) How may diversification of tree species within a wood allow the continued support of woodland biodiversity? 4) How do we implement and communicate management strategies to increase woodland resilience?

To address these knowledge gaps, we work across disciplines bringing together ecologists, microbiologists, social scientists, and woodland managers. The Woodland Trust is embedded at the heart of our project to enable us to co-develop and check the feasibility of our results with practitioners. Results from interviews with woodland managers, focus groups and analyses of policy documents, will be used to improve knowledge of the options for woodland diversification, and both the enthusiasm for, and capacity to, implement diversification strategies.

The microbes on leaves are important for plant health. Utilizing existing long-term experiments, we will examine the microbes on the leaves of Scots Pine grown in monocultures and in mixed woods. We will assess if the diversity of microbes on a leaf increases as the diversity of tree species increases, and whether this enables the trees to resist existing diseases.
Surprising we don't have lists of which species use which trees. This information is required if we are to plant trees that will continue to support woodland biodiversity. We will collate data on the biodiversity hosted by Scots Pine and Sitka Spruce and assess which other tree species could also support the same biodiversity.

Finally, we bring the results together to co-develop with practitioners, management strategies for diversification and case studies illustrating how the results can be implemented. The results will be shared via videos, podcasts, social media, and practitioner notes in addition to publications in the scientific literature.

Publications

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Dandy N (2024) The concept of woodland diversity: a review in Quarterly Journal of Forestry

 
Description We now know how many species use Scots pine and how many use Sitka spruce and if these species will use any of 46 other tree species that might be used to diversify Sitka spruce or Scots pine forests. We have shown that tree species diversification of Sitka spruce forests not only brings resilience for the biodiversity supported by Sitka spruce but provides support for a wide range of other biodiversity not hosted by Sitka spruce. However the tree species that bring the greatest biodiversity benefits will not grow in intimate mixes with Sitka spruce, so the planting design is critical. In contrast tree species diversification of Scots pine doesn't provide support for a lot of the biodiversity supported by Scots pine because many of the species supported by Scots pine are specialists.

We have found that woodland managers have a good understanding of diversification, across a range of managers with different objectives. However, in some cases managers felt that species diversification won't allow them to achieve their objectives be that production forestry or conservation. In addition the current grant system did not always help managers to diversify, it largely only supported them to do work they would have already done.

The diversity of the microbiome of trees leaves was shown to be more strongly influenced by the tree species present and the structure and age of the woodland, than the diversity of the tree species present.

We have identified the challenges associated with Continuous Cover Forestry, Natural Regeneration and enrichment planting, as methods to increase tree species diversity. We have shown that none of these methods will guarantee a more diverse forest in the long term, as this is dependent on the respective growth and shade tolerances of the tree species in the mix.
Exploitation Route It could be used in the identification of speices mixes to increase the resilience of forest associated biodiversity.
Sectors Environment

URL https://www.hutton.ac.uk/project/diversitree/
 
Description The findings have been used by CIEEM, the Royal Forestry Society, and the Community Woodlands association in their thinking around diversification. Specifically we have presented lunchtime webinars to all these groups. The results were also used in the Royal Society for Edinburgh report on Public financial support for tree planting and forestry https://rse.org.uk/programme/policy-advice/inquiries/tree-planting-inquiry/
First Year Of Impact 2024
Sector Environment
Impact Types Economic

Policy & public services

 
Description Attendance at Defra Trees and Woodlands Science Advisory meetings
Geographic Reach National 
Policy Influence Type Participation in a guidance/advisory committee
 
Description Contribution to RSE enquirey on Public financial support for tree planting and forestry
Geographic Reach National 
Policy Influence Type Contribution to a national consultation/review
URL https://rse.org.uk/expert-advice/inquiries/tree-planting-inquiry/
 
Description Invited to give evidence to the Rural Affairs and Islands committee of the Scottish Government on 15th January 2025
Geographic Reach National 
Policy Influence Type Participation in a guidance/advisory committee
URL https://www.parliament.scot/chamber-and-committees/official-report/search-what-was-said-in-parliamen...
 
Description RSE inquiry into public support for tree planting and forestry
Geographic Reach National 
Policy Influence Type Contribution to a national consultation/review
 
Description The Tree of Knowledge (ToK): communicating the complexity of forest resilience.
Amount £75,775 (GBP)
Funding ID NE/Y004124/1 
Organisation Natural Environment Research Council 
Sector Public
Country United Kingdom
Start 11/2023 
End 04/2025
 
Title DiversiTree 16S and ITS amplicon sequences 
Description DNA sequence of microbes isolated from trees in the UK 
Type Of Material Database/Collection of data 
Year Produced 2025 
Provided To Others? Yes  
Impact None as yet 
URL https://www.ncbi.nlm.nih.gov/bioproject/?term=PRJNA1183283
 
Title Scots pine-associated biodiversity in the UK (Scots pine Ecol) 
Description This is a database of all known birds, bryophytes, fungi, invertebrates, lichens and mammals that use Scots Pine (Pinus sylvestris) in the UK. Scots pine is the UK's only native conifer that is used commercially and is the most common tree in the UK's native Caledonian pine woods. In total 1589 species are listed in the database. For each species a level of association with Scots pine is provided, ranging from obligate (only found on Scots pine) to cosmopolitan (found on a wide range of other tree species). Data on the ecology of each Scots pine associated species were collated: part of tree used, and use made of tree (feeding, roosting, breeding). Data on use or otherwise of 34 other tree species were also collated for each of the 1589 species. 
Type Of Material Database/Collection of data 
Year Produced 2025 
Provided To Others? Yes  
Impact Ongoing paper writing, presentation of results to practitioners and discussion about implementation of results 
URL https://catalogue.ceh.ac.uk/id/97f8c19b-7451-4809-b011-0f9cf3eb8430
 
Title Sitka spruce-associated biodiversity in the UK (Sitka spruce Ecol) 
Description Here we collate a database of all known birds, bryophytes, fungi, invertebrates, lichens and mammals that use Sitka spruce (Picea sitchensis) in the UK. Sitka spruce is the most commonly planted commercial tree species in the UK. In total 564 species are listed in the database. For each species we provide a level of association with Sitka spruce, ranging from obligate (only found on Sitka spruce) to cosmopolitan (found on a wide range of other tree species). Data on the ecology of each Sitka spruce associated species were collated: part of tree used, and use made of tree (feeding, roosting, breeding). Data on use or otherwise of 34 other tree species were also collated for each of the 564 species. 
Type Of Material Database/Collection of data 
Year Produced 2024 
Provided To Others? Yes  
Impact Publications being prepared and talks given to practitioners resulting in discussion about diversification and future tree species. 
URL https://catalogue.ceh.ac.uk/id/1ce52c10-e3ab-4996-b9b7-052c70b3c1ba
 
Description DiversiTree - Diversifying our Woodlands to Increase Resilience A talk to Community Woodlands Association and LyG 
Form Of Engagement Activity A talk or presentation
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Professional Practitioners
Results and Impact A talk as a lunchtime webinar to the Community woodland association and their Welsh equivalent LyG to showcase the results from the project.
Year(s) Of Engagement Activity 2025
 
Description DiversiTree - Diversifying our Woodlands to Increase Resilience RFS webinar 
Form Of Engagement Activity A talk or presentation
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Professional Practitioners
Results and Impact A webinar hosted by the Royal Forestry society and presented to their members show casing the outcome of the project across all work packages. Around 70 people present.
Year(s) Of Engagement Activity 2025
 
Description DiversiTree - Diversifying our Woodlands to Increase Resilience Talk to CIEEM 
Form Of Engagement Activity A talk or presentation
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Professional Practitioners
Results and Impact A webinar hosted by CIEEM to show case the results of the project across the different work packages 248 people signed up
Year(s) Of Engagement Activity 2025
 
Description DiversiTree practitioners panel - Mar lodge 13 October 23 
Form Of Engagement Activity Participation in an activity, workshop or similar
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Professional Practitioners
Results and Impact A day visit to Mar lodge with forest practitioners to discuss natural regeneration, woodland expansion and forest divesification. We looked at the methods to do this and which factors were inhibitors. One participant said " I just wanted to say how much I enjoyed and valued the Mar Lodge visit and how wonderfully you looked after us all too!
It was perhaps the best field visit experience with the most wonderful and knowledgeable group of people that I can remember."
Year(s) Of Engagement Activity 2023
 
Description DiversiTree's practitioner pannel - a talk at the Treescapes final conference in June 24 
Form Of Engagement Activity A talk or presentation
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Professional Practitioners
Results and Impact A talk about how the project engaged with practitioners
Year(s) Of Engagement Activity 2024
 
Description Diversifying the tree species composition of Sitka spruce and Scots Pine forests to increase resilience for associated biodiversity. 
Form Of Engagement Activity A talk or presentation
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Professional Practitioners
Results and Impact A talk at the final Treescapes conference to which a range of policy representatives were present.
Year(s) Of Engagement Activity 2024
 
Description Diversifying tree species within our forests to increase resilience - an interdisciplinary approach. 
Form Of Engagement Activity A talk or presentation
Part Of Official Scheme? No
Geographic Reach Local
Primary Audience Other audiences
Results and Impact Talk at the Birmingham Institute of Forest Research Workshop
Year(s) Of Engagement Activity 2023
 
Description Diversifying tree species within our forests to increase resilience - an interdisciplinary approach. 
Form Of Engagement Activity A talk or presentation
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Other audiences
Results and Impact Talk at Hutton symposium
Year(s) Of Engagement Activity 2023
 
Description Future of UK Treescapes - public dialogue 
Form Of Engagement Activity A talk or presentation
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Public/other audiences
Results and Impact The Future of UK Treescapes programme organised a public consultation, with c 16 members of the public listening to online talks on Saturday 19th October. I gave a presentation on "Biodiversity and Treescapes" and answered questions
Year(s) Of Engagement Activity 2024
 
Description How much diversity is enough - a presentation to the final Treescapes Conference June 24 Glasgow 
Form Of Engagement Activity A talk or presentation
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Policymakers/politicians
Results and Impact A debate covering genetic, species and epigenetics and how much diversity is enough to provide resilience for future woods and forests
Year(s) Of Engagement Activity 2024
 
Description I grew up without Elm. What is my grandson growing up without? 
Form Of Engagement Activity Engagement focused website, blog or social media channel
Part Of Official Scheme? No
Geographic Reach International
Primary Audience Professional Practitioners
Results and Impact Blog outlining how increased tree diseases can have a cascading impact on biodiversity and lower our baseline biodiversity over generations. It discussed if greater diversity of tree species and greater biosecurity were part of the solution.
Year(s) Of Engagement Activity 2023
URL https://cieem.net/i-grew-up-without-elm-what-is-my-grandson-growing-up-without/
 
Description Podcast on visible and invisible forest diversity 
Form Of Engagement Activity Engagement focused website, blog or social media channel
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Public/other audiences
Results and Impact A podcast on forest diversity talking about the importance of species, genetic and epigenetic diversity and how this may influence resilience.
Year(s) Of Engagement Activity 2023
URL https://8point9.com/dr-ruth-mitchell-hutton-institute-visible-and-invisible-forest-diversity/
 
Description Practitioners Pannel to discuss diversity 
Form Of Engagement Activity Participation in an activity, workshop or similar
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Professional Practitioners
Results and Impact A meeting with practitioners to discuss the DiversiTree project, specifically to discuss different management techniques to increase tree species diversity. A mix of an indoor meeting and an outdoor meeting in the field. The event sparked lots of discussion over the pros and cons of increasing tree species diversity.
Year(s) Of Engagement Activity 2023
 
Description Presentation at AAB Creating Canopies conference on polarisation in the sector with DiversiTree PrP featured 
Form Of Engagement Activity A talk or presentation
Part Of Official Scheme? No
Geographic Reach International
Primary Audience Professional Practitioners
Results and Impact Presentation at the Association of Applied Biologists' Creating Canopies conference on polarisation in the forestry/conservation sector - using DiversiTree's Practitioner Panel as an example of bridging divides.
Year(s) Of Engagement Activity 2023
 
Description Prioritising Diversity? A Question for Woodland Managers Talk at Treescapes 24 final conference in Glasgow 
Form Of Engagement Activity A talk or presentation
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Professional Practitioners
Results and Impact Further discussion
Year(s) Of Engagement Activity 2024
 
Description Prioritising Diversity? A question for Woodland managers. Talk at Final Treescapes conference June 24 
Form Of Engagement Activity A talk or presentation
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Professional Practitioners
Results and Impact A talk summarizing the results from the social science work package on land managers perceptions of diversity and their opinions on the opportunities for and the barriers to diversification
Year(s) Of Engagement Activity 2024
 
Description Social media content 
Form Of Engagement Activity Engagement focused website, blog or social media channel
Part Of Official Scheme? No
Geographic Reach International
Primary Audience Professional Practitioners
Results and Impact Project social media channel content - Twitter
Year(s) Of Engagement Activity 2022,2023,2024
URL https://twitter.com/DiversiTree_UK
 
Description Talk to CIEEM 
Form Of Engagement Activity A talk or presentation
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Professional Practitioners
Results and Impact A webinar to CIEEM members describing the project. Provoked discussion around whether our woodlands are already diverse enough and which methods we might use to diversify them.
Year(s) Of Engagement Activity 2023
URL https://youtu.be/-_BBvFYzgHk
 
Description The role of the microbiome in tree resilience A talk at the final Treescapes conference in June 24 
Form Of Engagement Activity A talk or presentation
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Professional Practitioners
Results and Impact A talk on how the microbiome of the tree leaves changes between mono-cultures and mixed plantations and whether this infers resilience to tree diseases.
Year(s) Of Engagement Activity 2024
 
Description Treescapes Webinar 
Form Of Engagement Activity A talk or presentation
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Professional Practitioners
Results and Impact A webinar on the inital project outputs, with c20minutes discussion afterwards over a whole range of issues
Year(s) Of Engagement Activity 2024
 
Description Webinar to Chartered Institute of Foresters 
Form Of Engagement Activity A talk or presentation
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Professional Practitioners
Results and Impact A webinar describing the project with time for discussion on the subject of diversifying tree species composition. 129 participants.
Year(s) Of Engagement Activity 2023
 
Description Webinar to Royal Forestry Society 
Form Of Engagement Activity A talk or presentation
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Professional Practitioners
Results and Impact A webinar about the project with time for questions and discussion. 70 people present in total (although some logins had more than one person watching) and recorded for future use.
Year(s) Of Engagement Activity 2023
 
Description Webinar to Royal Forestry Society on 29th January 2025 
Form Of Engagement Activity A talk or presentation
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Professional Practitioners
Results and Impact A one hour webinar describing the results of the project with time for discussion. This resulted in an invitation to communicate the results more widely across RFS by writing an article in Quarterly Journal of Forestry.
Year(s) Of Engagement Activity 2025
 
Description Webinar to Small Woods Group 
Form Of Engagement Activity A talk or presentation
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Professional Practitioners
Results and Impact A webinar to woodland owners about the DiversiTree project
Year(s) Of Engagement Activity 2023
 
Description Webinar to Woodland Trust 
Form Of Engagement Activity A talk or presentation
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Professional Practitioners
Results and Impact A webinar for Woodland Trust staff describing the project and what we aimed to achieve
Year(s) Of Engagement Activity 2022
 
Description Workshop and Information Stall at Community Woodlands Association Annual Conference 
Form Of Engagement Activity Participation in an activity, workshop or similar
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Third sector organisations
Results and Impact Project members conducting a workshop at the CWA annual conference (part of data gathering for WP1) followed by a public facing stall to disseminate information about the project to the conference as a whole.
Year(s) Of Engagement Activity 2023
URL https://www.communitywoods.org/cwa-annual-conference-2023
 
Description Workshop and Information Stall at Community Woodlands Association Annual Conference 
Form Of Engagement Activity Participation in an activity, workshop or similar
Part Of Official Scheme? No
Geographic Reach National
Primary Audience Third sector organisations
Results and Impact Project members conducting a workshop at the CWA annual conference (part of data gathering for WP1) followed by a public facing stall to disseminate information about the project to the conference as a whole.
Year(s) Of Engagement Activity 2023
URL https://www.communitywoods.org/cwa-annual-conference-2023